When both partners benefit equally from a partnership, they are said to be in a symbiotic relationship. There are many such relationships in the natural world. For example, when a bee goes to a flower to collect nectar, it also brings about pollination by carrying pollen on its furry body from one flower to the next. Both the bee and the flower benefit.

A symbiosis is an evolved interaction or close living relationship between organisms from different species, usually with benefits to one or both of the individuals involved. Symbioses may be ‘obligate’, in which case the relationship between the two species is so interdependent, that each of the organisms is unable to survive without the other, or ‘facultative’, in which the two species engage in a symbiotic partnership through choice, and can survive individually. Obligate symbioses are often evolved over a long period of time, while facultative symbioses may be more modern, behavioral adaptions; given time, facultative symbioses may evolve into obligate symbioses.

Endosymbiosis is a symbiotic relationship, occurring when one of the symbiotic partners lives within the body of the other. Endosymbiosis can take place either within the cells (intercellular symbiosis) of the ‘host’ organism, or outside the cells (extracellular symbiosis). On the other hand, ectosymbiosis is a symbiotic relationship in which one organism lives on the body surface of the host, including the lining of the digestive tract, or exocrine glands such as mucus or sweat glands.

Mutualisms are a form of symbiosis in which both symbiotic partners benefit from the interaction, often resulting in a significant fitness gain for either one or both parties. Mutualisms can take the form of resource-resource relationships, service-resource relationships, or service-service relationships.

Resource-resource mutualisms (also known as ‘trophic mutualisms’) happen through the exchange of one resource for another between the two organisms involved. Resource-resource mutualisms most often occur between an autotroph (a photosynthesizing organism) and a heterotroph (an organism which must absorb or ingest food to gain energy). Most plants have a trophic mutualism called a mycorrhizal association, which is a symbiosis between the roots of the plants and a fungus. The fungus colonizes the plants roots and is provided with carbohydrates, sucrose and glucose. In exchange, the plant benefits from the fungi’s higher water and mineral absorption capabilities.

Service-resource mutualisms occur when the symbiotic partner provides a service in exchange for a resource reward. One of the best known examples of this is the exchange between plants and their pollinators. While visiting the plants to gain a supply of energy-rich nectar, the pollinator (insects, birds, moths, bats, etc.), provides the plant with the service benefit of being pollinated, while ensuring their own pollen is distributed when the pollinator visits more plants of the same species.

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